Five years after the occasions of Devil May Cry 4, Nero has set up his very own evil presence chasing organization based out of a van embellished with a neon "Fallen angel May Cry" sign Dante gave him with help from Kyrie and his specialist, Nico. On April 30, Nero experiences a perishing evil spirit who rips off his Devil Bringer arm and uses the Devil Arm Yamato to open an entry to escape in. A couple of days after the fact, a man known as "V" goes to the Devil May Cry office to employ Dante, Lady, and Trish to kill a specific returning evil presence. The gathering touch base in Red Grave City on May 16, running into Nero as they share a similar focus on: A devil named Urizen who has planted a satanic tree called Qliphoth in the city, which is murdering individuals for their blood. The gathered gathering endeavor to assault Urizen together, yet the evil spirit overwhelms them all; Dante is at last crushed and shot out from the Qliphoth with his sword Rebellion broke, Lady and Trish are caught and transformed into centers for the devils Artemis and Cavaliere Angelo, and V and Nero departure to security.
After a month on June 15, Nero comes back to Red Grave in the wake of being equipped with the "Fallen angel Breaker" prosthetic arm Nico made for him. Nero gets together with V subsequent to besting the evil presence Goliath, who is looking for Qliphoth for its natural product - conceived of consolidated human blood - which makes whomever expends it the ruler of the Underworld. As the pair crush the Qliphoth's underlying foundations while hunting down Dante, Nero massacres Artemis to save Lady while V kills the evil presence parasite Nidhogg, later supporting Nero against the root build Gilgamesh. V rejoins Nero subsequent to learning another evil spirit, Malphas, is additionally after the Qliphoth's organic product while looking for the Devil Sword Sparda in dread of its namesake's relatives procuring it. V divides from to discover the Sparda first and finds it alongside a sleeping Dante, whose nearness was being covered up by the sword. V stirs Dante by taking steps to wound him with the Sparda.
Subsequent to realizing what happened amid his month-long trance like state, Dante battles his approach to Urizen and liberates Trish from Cavaliere Angelo en route. V at that point uncovers to Trish that Urizen is really the evil presence side of Dante's sibling Vergil, who utilized Yamato's capacity to isolate his devil and human parts - the last of which showed as V himself, whose body is achieving its farthest point. Dante, having heard the full story when first gathering V, reasons Rebellion's capacity is the inverse - the capacity to bring together something - as he ventures to every part of the remainder of his youth home. Acknowledging why his dad gave him Rebellion, Dante wounds himself with the wrecked sword's grip to assimilate the Sparda into himself, obtaining his Sin Devil Trigger structure alongside his very own Devil Sword Dante.
As the Qliphoth falls, Nero leaves Red Grave City close by Nico while taking out any straggling evil spirits en route. Weeks after the fact, Trish and Lady are enlisted for another activity by Morrison, who Dante left responsible for his office before the occasions at Red Grave City. In the Underworld, Dante and Vergil duel each-other interminably - because of being similarly coordinated and the consistent impedance from evil spirits out to slaughter them - however their contention has all the earmarks of being on increasingly neighborly terms. The diversion finishes as Dante and Vergil battle against an evil spirit swarm with Dante saying his catchphrase "Bonanza!", a lot to Vergil's inconvenience.
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